- Rusty Beck (Graham Patrick Martin): Sharon Raydor's adoptive son, a former prostitute who witnessed Phillip Stroh burying a body. He has since started an investigative video blog called Identity and has started preparing for law school after finishing his journalism degree.
- Assistant Chief Russell Taylor (Robert Gossett): Assistant Chief of Operations until Season 5 episode "White Lies, Part 1". He was Captain Raydor's first immediate supervisor when she assumed command of the MCD. The former commanding officer of the Robbery-Homicide Division and later press liaison for the Major Crimes Division, Taylor long felt passed over and unappreciated.
- Deputy Chief Fritz Howard (Jon Tenney): Fritz Howard was a Special Agent for the F.B.I. and the liaison officer to the LAPD (The Closer and Major Crimes, Seasons 1 and 2). Upon his retirement, he is recruited to head the LAPD's Special Operations Bureau and assumes the rank of Deputy Chief (Major Crimes, Season 3 onward). He is currently the Acting Assistant Chief of Operations until a replacement for Chief Taylor can be found.
- Lieutenant Louie Provenza (GW Bailey): The division's senior member and Sharon's second-in-command. He is the immediate supervisor of the team in the field.
- Lieutenant Michael Tao (Michael Paul Chan). The squad's forensics and technology expert; Tao previously was assigned to the LAPD's Scientific Investigation Division (SID).
- Detective Julio Sanchez (Raymond Cruz). The squad's gang and weapons expert, experience gained from his days on the LAPD's gang intelligence unit.
- Detective Amy Sykes (Kearran Giovanni). Transferred from the Special Investigation Section (SIS) of Special Operations, she is an expert on undercover operations. Sykes is a military veteran who served in Afghanistan.
- Detective Wes Nolan (Daniel DiTomasso): Det. Nolan was transferred to the division after ending his SIS undercover assignment. He will be assigned to the MCD until he can finish testifying against the Z-Brotherhood.
- Civilian Surveillance Coordinator Buzz Watson (Phillip P. Keene). Buzz is the Scientific Investigation Division's civilian investigator assigned to the division. He is responsible for memorializing crime scenes, and managing technology in the field and in the squad's office. He also recently qualified as a LAPD reserve police officer.
- Dr. Fernando Morales (Jonathan Del Arco): A Los Angeles County Deputy Medical Examiner who coordinates with the team, conducting autopsies. Morales has a sharp medical mind and and equally sharp, sometimes sarcastic, wit. Morales is gay, and provides expertise on the gay community in Los Angeles.
- Investigator Kendall (Ransford Doherty): Kendall is responsible for examining the body at the crime scene and taking initial measurements, as well as preparing the body for transportation to the Los Angeles County Coroner's office.
- Deputy District Attorney Andrea Hobbs (Kathe Mazur): The DDA who works most frequently with Major Crimes, Hobbs is a team player.
- Deputy District Attorney Emma Rios (Nadine Velazquez): A new, inexperienced DDA under Hobbs' supervision, she is charged with preparing the case against Phillip Stroh (Season 2).
Recurring characters Edit
- Commander Ann McGinnis (Laurie Holden): Commander McGinnis is the second-in-command of the LAPD's Special Operations Bureau.
- Lieutenant Chuck Cooper (Malcolm-Jamal Warner): Lieutenant Cooper is the commanding officer of the Special Investigation Section (SIS) of the LAPD.
- Detective Robert Oderno (Brandon Barash): Robert Oderno is a detective in the LAPD's Missing Persons Unit.
- Gustavo Wallace (Rene Rosado): Gus Wallace is the older brother of murder victim Mariana Wallace. He is currently dating Rusty Beck.
- Deputy Chief Winnie Davis (Camryn Manheim): Chief Davis is a Deputy Chief of Operations for the LAPD and one of the candidates to replace Chief Taylor as the new Director of Operations (Season 5 winter episodes).
- Commander Leo Mason (Leonard Roberts): Commander Mason is the commanding officer of the Criminal Intelligence Division of the LAPD and one of the candidates to replace Chief Taylor as the new Director of Operations (Season 5 winter episodes).
Plot and Theme by SeasonEdit
The theme for season 1 is fairness. The season will explore how fair life can, or cannot be, both professionally and personally.
The theme for season 2 is identity.
The theme for season 3 is expectation.
The theme for season 4 is courage.
The theme for season 5 is balance.